FLOWERING BIOLOGY OF GALIUM ODORATUM (RUBIACEAE)
Vladimir N. Godin
Central Siberian Botanical Garden, SB RAS, Novosibirsk, Russia
Keywords: Galium odoratum, andromonoecy, flowering biology
Abstract
The anthecology of the perennial herbaceous long-rhizome polycarpic plant, Galium odoratum (L.) Scop., in the Moscow region was studied. The observations were carried out in 2021-2023 according to generally accepted methods. At low illumination (less than 15 % of total sunlight) only perfect flowers are formed. With increasing illumination (to up to 60 % of full sunlight in clear weather), both perfect and staminate flowers are formed on individuals. The perfect flowers of G. odoratum are strictly protandrous. The life span of perfect flowers was 12-14 days, that of staminate flowers up to 2 days. The opening of flowers in synflorescence and in its structural units (dichasia) occurs in a clear sequence following to the order of branching of the shoot on which the flowers are formed. According to the ratio of the number of pollen grains and ovules in perfect flowers (from 1358 to 3010), G. odoratum represents a facultatively xenogamous plant. In habitats with a high degree of illumination, the perfect and staminate flowers of G. odoratum are characterized by a morning opening rhythm with a maximum at 8 a.m. In low illumination, the flowers open a few hours later.
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